Zn++ dependent  acid  phosphatase from  chicken's  liver   was  purified     to  almost homogeneity by   heat treatment at 60T,   cation exchange chromatopaphy on CM-Cellulose at pH   5.5,  gel  filtration  on  Sephadex   G-150  and  affinity  chromatogmphy on Sepharosic tartramic acid  affinity gel with specific activity  of 4U/mg  and  recovery of  8%. Purification achieved   was   50  fold.   He   purified   enzyme migrated   a   single   band.  The  enzyme  ha molecular weight of    100,000 and is a dimer with   two apparent identical subunits of 48,000 - 50,000  as  determined  by  moditan dodecyl sulphate  polyacrylamide gel  electrophoresis. The enzyme requires Zn++   for catalytic activity. The maximum enzyme activity we obtained in the psesence of  5  -  8  mM  ZnCl2 at pH  6.0. Higher  concentrations of Zn++   were  found  to  be inhibitory. As compound  to Za++   activation, Mn++ and Co++   showed  32% and  16%  activation respectively. The enzyme had  optimum pH  5.5  -   7.5, optimum temperature   55°C and   was stable  at 40°C. The  enzyme showed restricted specificity. p nitrephenylphosphate we found to be    a   better   abstract while  phenylphosphate, 8-glyccrophosphate and  FMN    were   also hydrolyzed    at  a significant  rate. Other  substrates  like phosphoserine, ATP and  phosphoenol pyruvate etc.  were   not hydrolyzed. NaF,  a   strong inhibitor  of high  molecular weight   acid phosphatase, had no effect on Za++- dependent  acid phosphates activity. However, the enzyme was    inhibited   by   phosphate,  titrate,  EDTA   and    ATP.  The   Km   value    against  p- nitrophenylphosphate as a substrate, was found  to be  ImM  at pH  6.0. Phosphste and tartrate were   competitive inhibitors with Ki 0.5mM and  6mM respectively. ATP was noncompetitive inhibitor (Ki = 2.3mM) while EDTA was found to be uncompetitive inhibitor (Ki =2.5mM).


S A MALIK ,ASMA SAEED ,A SAEED ,